I am a tenure-track researcher/assistant professor at the Institute for Advanced Study in Toulouse, University of Toulouse 1 Capitole.

I am mainly interested in social evolution, with a focus on biological collective action problems.

I use mathematical and computational models from various theoretical frameworks to explore the strategic, demographic, and ecological factors influencing the evolution of social traits.

To know more about the header image (an excerpt from Max Ernst’s The Phases of the Night) see The Mathematics of Max Ernst.



jorge [dot] pena [at] iast [dot] fr

Current address

Institute for Advanced Study in Toulouse
Manufacture des Tabacs
office ME 503.4
21 allée de Brienne
31015 Toulouse Cedex 6


+33 5 61 12 86 18